1. Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran
2. Zoological Museum, Biodiversity Unit, FI-20014 University of Turku, Finland
3. A.O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nakhimov Ave. 2, Sevastopol 299011 Crimea, Russia
4. Department of Biology, Faculty of Science, Eskişehir Technical University, TR-26470 Eskişehir, Turkey