1. Altai State University, Lenina Pr., 61, Barnaul, 656049, Russia
2. Systematic Zoology Laboratory, Department of Biological Sciences, Tokyo Metropolitan University, 1-1 Minami-Osawa, Hachioji-shi, 192-0397, Tokyo, Japan
3. Institute for Biological Problems of the North of the Far Eastern Branch of the Russian Academy of Sciences, Portovaya str., 18, Magadan 685000 Russia.