1. Prioksko-Terrasnyi State Biosphere Reserve, Danki, Moscow Area 142200, Russia
2. Laboratory of Systematics of Invertebrate Animals, Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, Frunze street 11, Novosibirsk 630091, Russia
3. Department of Entomology, Faculty of Biology, Lomonosov Moscow State University, Leninskie Gory, Moscow 119234, Russia.