1. P.P.Shirshov Institute of Oceanology Russian Academy of Science, 36, Nakhimovsky prospect, Moscow, 117997, Russia
2. White Sea Biological Station, Department of Biology Lomonosov Moscow State University, P.O. Box 20, Glavpochtamt, Kandalaksha raion, Murmanskaya oblast 184042, Russia
3. Russian Federal Research Institute of Fisheries and Oceanography, 17 Verhnyaya Krasnoselskaya, Moscow 107140, Russia.