1. Marine Research Laboratory, Zoological Institute, Russian Academy of Sciences, Universitetskaya emb. 1, St. Petersburg 199034 Russia
2. Invertebrate Zoology Department, Lomonosov Moscow State University, Leninskie gori 1-12, Moscow 119234 Russia
3. A.V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch, Russian Academy of Sciences, Palchevskogo str. 17, Vladivostok 690041 Russia