1. The Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski pr. 33, Moscow 119071, Russia. E-mail: lsdc@mail.ru
2. Institute of Biology, Komi Science Center, Ural Division of Russian Academy of Sciences, Kommunisticheskaya str. 28, Syktyvkar 167982, Russia. E-mail: taskaeva@ib.komisc.ru