1. Botanical Garden-Institute FEB RAS, Makovskogo Street, 142, Vladivostok, 690024, Russia
2. Institute of Biology and Soil Science, Stoletiya Vladivostoka Avenue, 159, Vladivostok 690022 Russia
3. College of Life Sciences, Guizhou University, Guiyang, 550025, China
4. Institute of the Industrial Ecology Problems of the North, Kola Science Centre RAS, Fersman Str. 14a, Apatity, Murmansk Provinсe, 184209, Russia
5. Polar Alpine Botanical Garden and Institute, Kola Science Center RAS, Kirovsk-6, Murmansk Province 184256 Russia