1. Department of Watershed Sciences and the Ecology Center, Utah State University, 5210 Old Main Hill, Logan, Utah 84322-5210 USA, and Center for Ecological Research and Education, Department of Biological Sciences, Idaho State University, 921 S. 8th Ave., Stop 8007, Pocatello, Idaho 83209-8007 USA
2. Department of Biology and the Ecology Center, Utah State University, 5305 Old Main Hill, Logan, Utah 84322-5305 USA
3. Department of Watershed Sciences and the Ecology Center, Utah State University, 5210 Old Main Hill, Logan, Utah 84322-5210 USA