1. Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia 30602 USA
2. US Geological Survey, Georgia Cooperative Fish and Wildlife Research Unit, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia 30602 USA
3. Nongame Conservation Section, Wildlife Resources Division, Georgia Department of Natural Resources, Social Circle, Georgia 30025 USA
4. US Geological Survey, Southeast Ecological Research Center, School of Forest Resources and Conservation, University of Florida, Gainesville, Florida 32605 USA