1. Department of Forestry and Natural Resources, Clemson University, Clemson, South Carolina 29634-0317 USA
2. Department of Geography and Environmental Systems, University of Maryland, Baltimore County, Baltimore, Maryland 21250 USA
3. Department of Biology, Colorado State University, Fort Collins, Colorado 80523 USA
4. Northeast Amphibian Research and Monitoring Initiative, Patuxent Wildlife Center, US Geological Survey, Laurel, Maryland 20708 USA
5. US Geological Survey Leetown Science Center, Aquatic Ecology Branch, Kearneysville, West Virginia 25430 USA
6. US Geological Survey, Leetown Science Center, Northern Appalachian Research Laboratory, Wellsboro, Pennsylvania 16901 USA
7. Department of Biological Sciences, University of Notre Dame, Notre Dame, Indiana 46556 USA