1. a Department of Biology, Indiana State University, 200 North Seventh Street, Terre Haute, IN 47809, USA
2. b Mitrani Department of Desert Ecology, Jacob Blaustein Institutes for Desert Research, Ben-Gurion University of the Negev, Midreshet Ben-Gurion, 84990, Israel
3. c Conservation Science Department, Brookfield Zoo, Chicago Zoological Society, 3300 Golf Road, Brookfield, IL 60513, USA
4. d Department of Conservation and Research, Oregon Zoo, 4001 SW Canyon Road, Portland, OR 97221, USA