1. Ocean Tech Marine Institute, Mot-gol Beonyong-ro 71 beon-gil, Nam-gu, Busan 608-817, South Korea
2. Southeast Sea Fisheries Research Institute, National Fisheries Research and Development Institute, 397-68, Sanyangilju-ro, Sanyang-eup, Tongyeong 650-943, South Korea
3. Fisheries Resources Management Division, National Fisheries Research and Development Institute, 152-1, Haean-ro, Gijang-eup, Gijang-gun, Busan 619-705, South Korea
4. Korea Benthos Research Center, Co. Ltd., 14, Suyeong-ro 366 beon-gil, Nam-gu, Busan 608-804, South Korea