1. Department of Biological Sciences, University of Zimbabwe, P.O. Box MP167, Mount Pleasant, Harare, Zimbabwe
2. Zoology and Entomology, Rhodes University, P.O. Box 94, Grahamstown 6140, South Africa
3. Laboratory of Aquatic Ecology, Evolution and Conservation, KU Leuven, Charles Deberiotstraat 32, B-3000 Leuven, Belgium