1. 1Institute of Marine Biology, College of Life Sciences, National Taiwan Ocean University, 2 Pei-Ning Road, Keelung 20224, Taiwan
2. 2Department of Marine Science, Faculty of Fisheries, Kasetsart University, 50 Ngamwongwan Rd., Chatuchak, Bangkok 10900, Thailand
3. 3Department of Environment Biology and Fishery Science, National Taiwan Ocean University, 2 Pei-Ning Road, Keelung 20224, Taiwan
4. 4Center of Excellence for the Oceans, National Taiwan Ocean University, 2 Pei-Ning Road, Keelung 20224, Taiwan