1. 1Department of Aquaculture, Faculty of Agriculture, Universiti Putra Malaysia, 43400 UPM, Serdang, Selangor, Malaysia
2. 3International Institute of Aquaculture and Aquatic Sciences, Universiti Putra Malaysia, Jalan Kemang 6, Teluk Kemang 71050, Port Dickson, Malaysia
3. 2School of Marine Science and Technology, Tokai University, 3-20-1, Orido, Shimizu, Shizuoka, 424-8610 Japan