1. 1Graduate School of Biosphere Science, Hiroshima University, 1-4-4 Kagamiyama, Higashi-Hiroshima, Hiroshima 739-8528, Japan
2. 2Suzuka Branch, Mie Prefecture Fisheries Research Institute, 1-6277-4 Shiroko, Suzuka, Mie 510-0243, Japan
3. 3Graduate School of Science and Engineering, Kagoshima University, 1-21-35 Korimoto, Kagoshima, Kagoshima 890-0065, Japan