1. 1Doctoral Degree Program in Marine Biotechnology, National Sun Yat-sen University and Academia Sinica, Kaohsiung 80424, Taiwan
2. 2Biodiversity Research Center, Academia Sinica, Taipei 11529, Taiwan
3. 3Institute of Natural and Environmental Sciences, University of Hyogo, 6 Yayoigaoka, Sanda, Hyogo, 669-1546, Japan
4. 4Museum of Nature and Human Activities, Hyogo, 6 Yayoigaoka, Sanda, Hyogo, 669-1546, Japan
5. 5White Sea Biological Station, Biological Faculty, Moscow State University, 119991, Moscow, Russia