1. Doctoral School of Biology, Institute of Biology, ELTE Eötvös Loránd University, Pázmány Péter sétány 1C, Budapest 1117, Hungary
2. Department of Systematic Zoology and Ecology, Eötvös Loránd University, Pázmány Péter sétány 1C, Budapest 1117, Hungary
3. Department of Zoology, Swedish Museum of Natural History, Box 50007, SE-104 05 Stockholm, Sweden