1. Department of Natural Science, Elizabeth City State University, Elizabeth City, NC 27909, USA
2. Department of Entomology and Nematology, University of Florida, Gainesville, FL 32611, USA
3. Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000, Brussels, Belgium
4. Biology Department, Ghent University, Ledeganckstraat 35, 9000 Ghent, Belgium