1. Far Eastern Federal University, 27 Oktyabrskaya Street, Vladivostok 690091, Russia
2. Institute of Biology and Soil Science FEB RAS, 159 Prospect 100-letija, Vladivostok 690022, Russia
3. A.V. Zhirmunsky Institute of Marine Biology FEB RAS, 17 Paltchevsky Street, Vladivostok 690041, Russia