1. 1Southern Research Station, USDA Forest Service Athens, GA 30602, USA
2. 2Forest Health Protection, USDA Forest Service, Athens, GA 30602, USA
3. 3USDA Agricultural Research Service, Nematology Laboratory, BARC-West, Beltsville, MD 20705, USA
4. 4Southern Research Station, USDA Forest Service, Asheville, NC 28804, USA