1. 1School of Biological Sciences, Brambell Building, University of Wales, Bangor LL57 2UW, United Kingdom
2. 2Department of Zoology, University of Otago, P.O. Box 56, Dunedin, New Zealand
3. 3School of Biological Sciences, Brambell Building, University of Wales, Bangor LL57 2UW, United Kingdom
4. 4Department of Ecology, Faculty of Biology, University of Bucharest, Splaiul Independentei 91-95, 76201 Bucharest, Romania