1. 1Museo de Ciencias Naturales de Tenerife, Apdo. 853, 38080 Santa Cruz de Tenerife, Islas Canarias, Spain
2. 2Muséum National d' Histoire Naturelle, Laboratoire de Paléontologie, URA 12- CNRS, 8 rue Buffon, 75005 Paris, France
3. 3Department of Anatomy and Developmental Biology, University College London, Rockefeller Building, University Street, London WC1E 6JJ, Great Britain