1. 1Entomology and Nematology Department, IACR-Rothamsted, Harpenden, Herts, AL5 2JQ and Department of Biology, Imperial College of Science, Technology and Medecine, Silwood Park, Ascot, Berkshire, SL5 7PY, UK
2. 2Department of Biology, Imperial College of Science, Technology and Medecine, Silwood Park, Ascot, Berkshire, SL5 7PY, UK
3. 3Entomology and Nematology Department, IACR-Rothamsted, Harpenden, Herts, AL5 2JQ