1. a Zoological Institute of the Russian Academy of Sciences, Universitetskaya emb., 1, St Petersburg, 199034, Russia
2. b Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan, 367000, Russia
3. c Dagestan State University, M. Gadzhiev str. 43a, Makhachkala, Republic of Dagestan, 367000, Russia
4. d Ege University, Zoology Department, Bornova–Izmir, 35100 Turkey Version of Record, published online 1 September 2019; published in print 10 August 2020