1. 1Ethology and Socio-ecology, University of Utrecht, P.O. Box 80086, 3508 TB Utrecht, The Netherlands
2. 2Ethology and Socio-ecology, University of Utrecht, P.O. Box 80086, 3508 TB Utrecht, The Netherlands
3. 3Ethology and Socio-ecology, University of Utrecht, P.O. Box 80086, 3508 TB Utrecht, The Netherlands
4. 4Ethology and Socio-ecology, University of Utrecht, P.O. Box 80086, 3508 TB Utrecht, The Netherlands