1. 1Marine Biology Laboratories, 7 MBL Street, Woods Hole, MA 02543, Institut für Verhaltensbiologie, Freie Universität Berlin, Haderslebenstrasse 9, 12163 Berlin, Germany
2. 2Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544-1003, USA
3. 3Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544-1003, USA, Department of Biology, University of Leeds, Leeds LS2 9JT, UK