1. 1Department of Zoology, University of Uppsala, Villav. 9, S-752 36 Uppsala, Sweden, Department of Zoology, University of Basel, Rheinsprung 9, CH-4051 Basel, Switzerland
2. 2Department of Zoology, University of Uppsala, Villav. 9, S-752 36 Uppsala, Sweden;, Email: Anders.Berglund@zoologi.uu.se
3. 3Department of Zoology, Norwegian University of Technology and Sciences, Trondheim, N-7055 Dragvoll, Norway;, Email: gunros@alfa.avh.unit.no