1. Wildlife Management Center, Tokyo University of Agriculture and Technology, 3-5-8 Saiwai-Cho, Fuchu, Tokyo 183-8509, Japan
2. Mount Fuji Research Institute, Yamanashi Prefecture Government, 5597-1 Kenmarubi, Kamiyoshida, Fijiyoshida, Yamanashi 403-0005, Japan
3. Laboratory of Wildlife Ecology and Conservation, School of Veterinary Medicine, Azabu University, 1-17-71 Fuchinobe, Chuo-ku, Sagamihara, Kanagawa 252-5201, Japan
4. Mount Asama Japanese Serow Research Group, 2-40 Saiwai-Cho, Fuchu, Tokyo 183-0054, Japan
5. Department of Forest Science, Faculty of Regional Environmental Science, Tokyo University of Agriculture, 1-1-1 Sakuragaoka, Setagaya 156-8502, Japan