1. aDivision of Biology, Graduate School of Science and Engineering, Yamagata University, 990-8560 Yamagata, Japan
2. bTokyo University of Marine Science and Technology, 4-5-7 Konan, Minato, 108-8477 Tokyo, Japan
3. cDepartment of Biology, Faculty of Science, Yamagata University, 990-8560 Yamagata, Japan