1. Department of Evolutionary Studies of Biosystems, The Graduate University for Advanced Studies, SOKENDAI, Shonan Village, Hayama, Miura, Kanagawa 240-0193, Japan
2. Department of Ecology and Evolutionary Biology, University of California, Los Angeles, 621 Young Drive South, Los Angeles, CA 90095-1606, USA
3. Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1, Yayoi, Bunkyo, Tokyo 113-8657, Japan
4. Department of Veterinary Nursing and Technology, Nippon Veterinary and Life Science University, 1-7-1 Kyonan-cho, Musashino, Tokyo 180-8602, Japan
5. Research Center for Integrative Evolutionary Science, The Graduate University for Advanced Studies, SOKENDAI, Shonan Village, Hayama, Miura, Kanagawa 240-0193, Japan