1. Deparment of Ecology and Evolution, Estación Biológica de Doñana (CSIC), 41092 Sevilla, Spain
2. Division of Behavioural Ecology, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland
3. Osa Conservation, Conservation Science Team, Washington, DC, 20005, USA
4. Remote Sensing and GIS Laboratory (LAST-EBD), Estación Biológica de Doñana (CSIC), 41092 Sevilla, Spain
5. Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, G12 8QQ, UK