1. Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand, arthitpolyotha@gmail.com
2. Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand, jirasak4@yahoo.com
3. Fauna and Flora International, Sanchaung Township, Yangon, Myanmar, aunglin199@gmail.com
4. Corresponding author; Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand
5. Academy of Science, The Royal Society of Thailand, Bangkok 10330, Thailand, somsak.pan@chula.ac.th