1. Department of Invertebrate Zoology, Biological Faculty, Lomonosov Moscow State University, Moscow, Russia
2. Taxonomy and Systematics Group, Naturalis Biodiversity Center, Leiden, The Netherlands
3. Groningen Institute for Evolutionary Life Sciences, University of Groningen, Groningen, The Netherlands
4. Taxonomy and Systematics Group, Naturalis Biodiversity Center, Leiden, The Netherlands, ivanenko.slava@gmail.com