1. Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Center for Morphology and Taxonomy, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. E-mail:
2. National Institute of Biological Resources, Biological and Genetic Resources Utilization Division, Environmental Research Complex, Hwangyeong-ro 42, Seo-gu, Incheon, 22689 Republic of Korea
3. Western Australian Museum, Collections & Research, 49 Kew Street, Welshpool, Western Australia 6106, Australia