1. Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, D-10115 Berlin, Germany; E-mail:
2. Department of Animal and Plant Sciences, University of Sheffield, Sheffield S10 2TN, U.K.
3. Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024-5192, U.S.A.
4. Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943, U.S.A., current address: 1950 Price Drive, Farmville, VA 23901, U.S.A.