1. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, The Chinese Academy of Sciences, Wuhan 430074, PR China
2. Institute of Botany, The Chinese Academy of Sciences, Beijing 100093, PR China
3. Graduate School of Chinese Academy of Sciences, Beijing, 100049, PR China
4. Key Laboratory of Horticultural Plant Biology, College of Horticulture and Forestry College, Huazhong Agricultural University, Wuhan 430070, PR China