1. Department of Agricultural, Food and Nutritional Science, University of Alberta, Edmonton, Alberta, Canada T6G 2P5
2. Alberta Environment and Sustainable Resource Development, Edmonton, Alberta, Canada T5K 2J6
3. Department of Renewable Resources, University of Alberta, Edmonton, Alberta, Canada T6G 2H1
4. Range Branch, Ministry of Forests, Lands and Natural Resource Operations, Prince George, British Columbia, Canada V2N 4W5