1. University of Alberta, Earth and Atmospheric Sciences. Edmonton, Alberta, Canada T6G 2E3
2. University of Alberta, Department of Biological Sciences, Edmonton, Alberta, Canada T6G 2E9
3. Cold Regions Research Centre, Wilfrid Laurier University, Waterloo, Ontario, Canada N2L 3C5
4. CSIRO Land and Water, Glen Osmond, South Australia, Australia SA 5064