1. RIKEN BioResource Research Center, Ibaraki 305-0074, Japan
2. Laboratory of Animal Reproduction, Graduate School of Veterinary Science, Azabu University, Kanagawa 252-5201, Japan
3. RIKEN Cluster for Pioneering Research, Saitama 351-0198, Japan
4. Graduate School of Life and Environmental Sciences, University of Tsukuba, Ibaraki 305-8572, Japan