1. RIKEN BioResource Center, Ibaraki 305-0074, Japan
2. Department of Life Science, School of Agriculture, Meiji University, Kanagawa 214-8571, Japan
3. Graduate School of Life and Environmental Science, University of Tsukuba, Ibaraki 305-8572, Japan
4. The Center for Disease Biology and Integrative Medicine, Faculty of Medicine, University of Tokyo, Tokyo 113-0033, Japan