1. Advanced Biotechnology Center, University of Yamanashi, Yamanashi 400-8510, Japan
2. Faculty of Life and Environmental Sciences, University of Yamanashi, Yamanashi 400-8510, Japan
3. RIKEN BioResource Research Center, Ibaraki 305-0074, Japan
4. RIKEN Cluster for Pioneering Research, Saitama 351-0198, Japan
5. Graduate School of Science and Technology, University of Tsukuba, Ibaraki 305-8577, Japan