1. Institute of Infection and Global Health, University of Liverpool; Leahurst Campus, Chester High Road, Neston CH64 7TE UK
2. NIHR Health Protection Research Unit in Emerging and Zoonotic Infections; The Farr Institute@HeRC, University of Liverpool; Waterhouse Building Liverpool L69 3GL UK
3. Bacteriology Department; APHA Weybridge; Addlestone Surrey KT15 3NB UK