1. Department of Clinical Veterinary Science; University of Bristol; Langford House, Langford Bristol BS40 5DU
2. Department of Pathology and Microbiology; University of Bristol; Langford House, Langford Bristol BS40 5DU
3. Faculty of Agricultural and Food SciencesM; University of Nottingham; Sutton Bonington Loughborough LE12 5RD