1. Department of Veterinary Clinical Sciences; Royal Veterinary College, Hawkshead Lane Hatfield Hertfordshire AL97TA UK
2. Department of Pathology and Pathogen Biology; Royal Veterinary College, Hawkshead Lane Hatfield Hertfordshire AL97TA UK
3. Institute of Veterinary; Animal and Biomedical Sciences; Massey University; Private Bag 11-222 Palmerston North New Zealand