1. Scottish Centre for Production Animal Health and Food Safety, School of Veterinary Medicine, University of Glasgow; Bearsden Road Glasgow G61 1QH UK
2. Moredun Scientific; Pentlands Science Park, Bush Loan Penicuik Midlothian EH26 0PZ UK
3. Veterinary Genes and Proteins Group; Institute of Biodiversity, Animal Health and Comparative Medicine, Garscube Campus, Glasgow University; Bearsden Road Glasgow G61 1QH UK