1. King Khalid Wildlife Research Centre, Thumamah; National Commission for Wildlife Conservation and Development; PO Box 61681 Riyadh 11575 Saudi Arabia
2. School of Life Sciences, Faculty of Science; Kingston University; Kingston upon Thames Surrey KT1 2EE
3. Department of Zoology, College of Science; King Saud University; PO Box 2455 Riyadh 11451 Saudi Arabia