1. Lambert; Leonard & May, Old Woodhouses, Broughall Whitchurch Shropshire SY13 4AQ UK
2. School of Veterinary Science; University of Liverpool; Leahurst Neston Cheshire CH64 7TE UK
3. Institute of Infection and Global Health; University of Liverpool; Leahurst Neston Cheshire CH64 7TE UK
4. Division of Animal Sciences; School of Biosciences; University of Nottingham; Sutton Bonington Campus Loughborough LE12 5RD UK