1. Department of Infection Biology; Institute of Infection and Global Health, School of Veterinary Science, University of Liverpool; Science Park ic2, 146 Brownlow Hill Liverpool Merseyside L3 5RF UK
2. Lanes Veterinary Group; Green Lane Veterinary Centre; Leachfield Industrial Estate, Green Lane West Cabus Garstang PR3 1PR UK
3. Department of Epidemiology and Population Health; Institute of Infection and Global Health, University of Liverpool; Leahurst Campus Neston Wirral CH64 7TE UK