1. Department of Molecular and Cellular Biology, College of Biological Science, University of Guelph, Guelph, Ontario, N1G 2W1, Canada.
2. Department of Plant Breeding and Genetics, University of Agriculture Faisalabad, Pakistan.
3. Department of Plant Agriculture, University of Guelph, Guelph, Ontario, N1G 2W1, Canada.
4. Agriculture and Agri-Food Canada, Food Research Program, 93 Stone Road West, Guelph, Ontario, N1G 5C9, Canada.